JOINT FILING AGREEMENT The undersigned, MVIL, LLC is jointly filing the attached Form 4 with MORNINGSIDE VENTURE INVESTMENTS LIMITED with respect to the beneficial ownership of securities of GreenLight Biosciences Holdings, PBC. For and on behalf of...

                             JOINT FILING AGREEMENT

The undersigned, MVIL, LLC is jointly filing the attached Form 4 with
MORNINGSIDE VENTURE INVESTMENTS LIMITED with respect to the beneficial ownership
of securities of GreenLight Biosciences Holdings, PBC.

For and on behalf of

MVIL, LLC


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▇▇▇▇▇ ▇▇▇ ▇▇▇▇ ▇▇▇▇▇, Manager